Convention relative à la conservation

de la vie sauvage et du milieu naturel de l'Europe

Comité permanent

Résolution n° 4 (1996) dressant l'inventaire des habitats naturels menacés nécessitant des mesures de conservation spécifiques

(adoptée par le Comité permanent le 6 décembre 1996)

Le Comité permanent de la Convention relative à la conservation de la vie sauvage et du milieu naturel de l'Europe, agissant en vertu de l'article 14 de la convention,

Eu égard à sa Résolution n° 1 (1989) concernant les dispositions relatives à la protection des habitats ;

Eu égard à sa Recommandation n° 14 (1989) concernant la conservation des habitats des espèces et la conservation des habitats naturels menacés ;

Reconnaissant que, pour les Parties Contractantes qui sont Etats membres de l'Union européenne, la liste des habitats naturels nécessitant des mesures de conservation spécifiques correspond à l'Annexe I de la Directive du Conseil 92/43/CEE du 21 mai 1992 concernant la conservation des habitats naturels ainsi que de la faune et de la flore sauvages,

Décide de conférer aux types d'habitats naturels énumérés dans l'annexe I de la présente Résolution le statut d'habitats naturels menacés nécessitant des mesures spécifiques de conservation (les unités élémentaires sont affectées du symbole !) ;

Décide de remettre régulièrement à jour l'annexe I de cette Résolution.

Revised Annex I of Resolution 4 (1996) of the Bern Convention

on endangered natural habitat types using the EUNIS habitat classification

(Adopted by the Standing Committee on 9 December 2010)

ENDANGERED NATURAL HABITAT TYPES

     A               Marine habitats

     A1             Littoral rock and other hard substrata

     A1.1          High energy littoral rock

!    A1.11        Mussel and/or barnacle communities

     A4.14        Mediterranean and Black Sea communities of lower mediolittoral rock very exposed to wave action

!    A1.141      Association with [Lithophyllum byssoides]

     A1.2          Moderate energy littoral rock

!    A1.22        Mussels and fucoids on moderately exposed shores

     A1.4          Features of littoral rock

!    A1.44        Communities of littoral caves and overhangs

     A2             Littoral sediment

!    A2.2          Littoral sand and muddy sand

!    A2.3          Littoral mud

!    A2.4          Littoral mixed sediments

!    A2.5          Coastal saltmarshes and saline reedbeds

                      includes the following subtypes separately listed in or split units from the 1998 version:

A2.521      Atlantic and Baltic brackish saltmarsh communities

A2.531      Atlantic upper shore communities

A2.542      Atlantic lower shore communities

A2.5514    [Salicornia veneta] swards

A2.5515    Black Sea annual [Salicornia], [Suaeda] and [Salsola] saltmarshes

A2.553      Atlantic [Sagina maritima] communities

     A2.6          Littoral sediments dominated by aquatic angiosperms

!   A2.61        Seagrass beds on littoral sediments

!   A2.621      [Eleocharis] beds

     A2.7          Littoral biogenic reefs

!    A2.72        Littoral mussel beds on sediment

!    A3             Infralittoral rock and other hard substrata

                      includes the following subtypes separately listed in or split units from the 1998 version:

A3.71        Robust faunal cushions and crusts in surge gullies and caves

A3.74        Caves and overhangs in infralittoral rock

!    A4             Circalittoral rock and other hard substrata

                       includes the following subtypes separately listed in or split units from the 1998 version:

A4.24        Mussel beds on circalittoral rock

A4.26        Mediterranean coralligenous communities moderately exposed to hydrodynamic action

A4.32        Mediterranean coralligenous communities sheltered from hydrodynamic action

A4.71        Communities of circalittoral caves and overhangs

!    A5             Sublittoral sediment

                      includes the following subtypes separately listed in or split units from the 1998 version:

A5.627      Baltic mussel beds in the infralittoral photic zone

     A6             Deep-sea bed

     A6.9          Vents, seeps, hypoxic and anoxic habitats of the deep sea

       A6.91        Deep-sea reducing habitats

!    A6.911      Seeps in the deep-sea bed

     B               Coastal habitats

     B1             Coastal dunes and sandy shores

!    B1.3          Shifting coastal dunes

!    B1.4          Coastal stable dune grassland (grey dunes)

!    B1.5          Coastal dune heaths

!    B1.6          Coastal dune scrub

!    B1.7          Coastal dune woods

!    B1.8          Moist and wet dune slacks

!    B1.9          Machair

     B2             Coastal shingle

!    B2.3          Upper shingle beaches with open vegetation

     C               Inland surface waters

     C1             Surface standing waters

!    C1.1          Permanent oligotrophic lakes, ponds and pools

                       includes the following subtype separately listed in or split unit from the 1998 version:

C1.14         Charophyte submerged carpets in oligotrophic waterbodies

     C1.2          Permanent mesotrophic lakes, ponds and pools

     C1.22        Free-floating vegetation of mesotrophic waterbodies

!    C1.222      Floating [Hydrocharis morsus-ranae] rafts

!    C1.223      Floating [Stratiotes aloides] rafts

!    C1.224      Floating [Utricularia australis] and [Utricularia vulgaris] colonies

!    C1.225      Floating [Salvinia natans] mats

!    C1.226      Floating [Aldrovanda vesiculosa] communities

     C1.24        Rooted floating vegetation of mesotrophic waterbodies

     C1.241      Floating broad-leaved carpets

!    C1.2416    [Nelumbo nucifera] beds

!    C1.25        Charophyte submerged carpets in mesotrophic waterbodies

     C1.3          Permanent eutrophic lakes, ponds and pools

     C1.34        Rooted floating vegetation of eutrophic waterbodies

     C1.341      Shallow-water floating communities

!    C1.3411    [Ranunculus] communities in shallow water

!    C1.3413    [Hottonia palustris] beds in shallow water

     C1.4          Permanent dystrophic lakes, ponds and pools

!    C1.44        Charophyte submerged carpets in dystrophic waterbodies

!    C1.5          Permanent inland saline and brackish lakes, ponds and pools

     C1.6          Temporary lakes, ponds and pools

!    C1.66        Temporary inland saline and brackish waters

!    C1.67        Turlough and lake-bottom meadows

     C2             Surface running waters

     C2.1          Springs, spring brooks and geysers

!    C2.12        Hard water springs

     C3             Littoral zone of inland surface waterbodies

     C3.4          Species-poor beds of low-growing water-fringing or amphibious vegetation

!    C3.41        Euro-Siberian perennial amphibious communities

     C3.42        Mediterraneo-Atlantic amphibious communities

!    C3.421      Short Mediterranean amphibious communities

!    C3.422      Tall Mediterranean amphibious communities

     C3.43        Central Eurasian amphibious communities

!    C3.431      Ponto-Pannonic riverbank dwarf sedge communities

     C3.5          Periodically inundated shores with pioneer and ephemeral vegetation

     C3.51        Euro-Siberian dwarf annual amphibious swards

!    C3.511      Freshwater dwarf [Eleocharis] communities

!    C3.512      Dune-slack [Centaurium] swards

!    C3.5132    Swards of small [Cyperus] species

!    C3.5133    Wet ground dwarf herb communities

!    C3.55        Sparsely vegetated river gravel banks

     C3.6          Unvegetated or sparsely vegetated shores with soft or mobile sediments

!    C3.62        Unvegetated river gravel banks

     D               Mires, bogs and fens

     D1             Raised and blanket bogs

!    D1.2          Blanket bogs

     D2             Valley mires, poor fens and transition mires

     D2.2          Poor fens and soft-water spring mires

     D2.22        [Carex nigra], [Carex canescens], [Carex echinata] fens

!    D2.226      Peri-Danubian black-white-star sedge fens

!    D2.3          Transition mires and quaking bogs

                       includes the following subtype separately listed in or split unit from the 1998 version:

D2.3H     Wet, open, acid peat and sand, with [Rhynchospora alba] and [Drosera]

     D3             Aapa, palsa and polygon mires

!    D3.1          Palsa mires

!    D3.2          Aapa mires

!    D3.3          Polygon mires

     D4             Base-rich fens and calcareous spring mires

!    D4.1          Rich fens, including eutrophic tall-herb fens and calcareous flushes and soaks

!    D4.2          Basic mountain flushes and streamsides, with a rich arctic-montane flora

     D5             Sedge and reedbeds, normally without free-standing water

!    D5.2          Beds of large sedges normally without free-standing water

     D6             Inland saline and brackish marshes and reedbeds

!    D6.1          Inland saltmarshes

                      includes the following subtypes separately listed in or split units from the 1998 version:

D6.15     Interior Iberian [Microcnemum] and [Salicornia] swards

D6.16     Interior central European and Anatolian [Salicornia], [Microcnemum], [Suaeda] and [Salsola] swards

     E                Grasslands and lands dominated by forbs, mosses or lichens

     E1              Dry grasslands

     E1.1           Inland sand and rock with open vegetation

     E1.11         Euro-Siberian rock debris swards

!    E1.112       [Sempervivum] or [Jovibarba] communities on rock debris

!    E1.2           Perennial calcareous grassland and basic steppes

!    E1.3           Mediterranean xeric grassland

     E1.7           Closed non-Mediterranean dry acid and neutral grassland

!    E1.71         [Nardus stricta] swards

     E1.8           Closed Mediterranean dry acid and neutral grassland

!    E1.83         Mediterraneo-montane [Nardus stricta] swards

!    E1.B          Heavy-metal grassland

     E2              Mesic grasslands

     E2.2           Low and medium altitude hay meadows

!    E2.25         Continental meadows

     E3              Seasonally wet and wet grasslands

!    E3.1           Mediterranean tall humid grassland

                      includes the following subtypes separately listed in or split units from the 1998 version:

E3.111    [Serapias] grassland

!    E3.4           Moist or wet eutrophic and mesotrophic grassland

!    E3.5           Moist or wet oligotrophic grassland

     E5              Woodland fringes and clearings and tall forb stands

     E5.4           Moist or wet tall-herb and fern fringes and meadows

     E5.41         Screens or veils of perennial tall herbs lining watercourses

     E5.411       Watercourse veils (other than of [Filipendula])

!    E5.4111     [Angelica archangelica] fluvial communities

!    E5.4112     [Angelica heterocarpa] fluvial communities

!    E5.4113     [Althaea officinalis] screens

!    E5.414       Continental river bank tall-herb communities dominated by [Filipendula]

!    E5.415       Eastern nemoral riverbanks with tall herb communities

     E5.42         Tall-herb communities of humid meadows

!    E5.423       Continental tall-herb communities of humid meadows

!    E5.424       Eastern nemoral Tall-herb communities of humid meadows

     E6              Inland salt steppes

!    E6.1           Mediterranean inland salt steppes

!    E6.2           Continental inland salt steppes

                       includes the following subtype separately listed in or split unit from the 1998 version:

E6.23      Central Eurasian solonchak grassland with [Crypsis]

     E7              Sparsely wooded grasslands

!    E7.3           Dehesa

     F                Heathland, scrub and tundra

     F2              Arctic, alpine and subalpine scrub

     F2.2           Evergreen alpine and subalpine heath and scrub

     F2.22         Alpide acidocline [Rhododendron] heaths

!    F2.224       Carpathian [Rhododendron kotschyi] heaths

!    F2.225       Balkan [Rhododendron kotschyi] heaths

!    F2.26         [Bruckenthalia] heaths

     F3              Temperate and mediterranean-montane scrub

     F3.2           Submediterranean deciduous thickets and brushes

     F3.24         Subcontinental and continental deciduous thickets

!    F3.241       Central European subcontinental thickets

     F4              Temperate shrub heathland

!    F4.1           Wet heaths

!    F4.2           Dry heaths

!    F4.3           Macaronesian heaths

     F5              Maquis, arborescent matorral and thermo-Mediterranean brushes

     F5.5           Thermo-Mediterranean scrub

!    F5.52         [Euphorbia dendroides] formations

!    F5.54         [Chamaerops humilis] brush

!    F5.55         Mediterranean pre-desert scrub

!    F5.56         Thermo-Mediterranean broom fields (retamares)

!    F5.5B        Cabo de Sao Vicente brushes

     F6              Garrigue

!    F6.7           Mediterranean gypsum scrubs

!    F6.8           Xero-halophile scrubs

!    F7              Spiny Mediterranean heaths (phrygana, hedgehog-heaths and related coastal cliff vegetation)

     F9              Riverine and fen scrubs

!    F9.1           Riverine scrub

!    F9.3          Southern riparian galleries and thickets (Excluding F9.35: Riperian stands of invasive shrubs)

     G               Woodland, forest and other wooded land

     G1             Broadleaved deciduous woodland

     G1.1          Riparian and gallery woodland, with dominant [Alnus], [Betula], [Populus] or [Salix]

!    G1.11        Riverine [Salix] woodland

!    G1.12        Boreo-alpine riparian galleries

!    G1.13        Southern [Alnus] and [Betula] galleries

     G1.2          Mixed riparian floodplain and gallery woodland

!    G1.21        Riverine [Fraxinus] - [Alnus] woodland, wet at high but not at low water

     G1.22        Mixed [Quercus] - [Ulmus] - [Fraxinus] woodland of great rivers

!    G1.221      Great medio-European fluvial forests

!    G1.223      Southeast European [Fraxinus] - [Quercus] - [Alnus] forests

!    G1.224      Po [Quercus] - [Fraxinus] - [Alnus] forests

     G1.3          Mediterranean riparian woodland

!    G1.36        Ponto-Sarmatic mixed [Populus] riverine forests

!    G1.37        Irano-Anatolian mixed riverine forests

!    G1.38        [Platanus orientalis] woods

!    G1.39        [Liquidambar orientalis] woods

     G1.4          Broadleaved swamp woodland not on acid peat

     G1.41        [Alnus] swamp woods not on acid peat

     G1.411      Meso-eutrophic swamp alder woods

!    G1.4115    Eastern Carpathian [Alnus glutinosa] swamp woods

!    G1.414      Steppe swamp [Alnus glutinosa] woods

!    G1.44        Wet-ground woodland of the Black and Caspian Seas

     G1.5          Broadleaved swamp woodland on acid peat

!    G1.51        Sphagnum [Betula] woods

!    G1.6          [Fagus] woodland

!    G1.7          Thermophilous deciduous woodland (excluding G1.7D Castanea sativa woodland)

                       includes the following subtypes separately listed in or split units from the 1998 version:

G1.7B     [Quercus pyrenaica] woodland

G1.7C     Mixed thermophilous woodland

!    G1.8          Acidophilous [Quercus]-dominated woodland

     G1.A         Meso- and eutrophic [Quercus], [Carpinus], [Fraxinus], [Acer], [Tilia], [Ulmus] and related woodland

!    G1.A1       [Quercus] - [Fraxinus] - [Carpinus betulus] woodland on eutrophic and mesotrophic soils

!    G1.A4       Ravine and slope woodland

!    G1.A7       Mixed deciduous woodland of the Black and Caspian Seas

!    G2             Broadleaved evergreen woodland (excluding G2.8 Highly artificial broadleaved evergreen forestry plantations and G2.9 Evergreen orchards and groves)

     G3             Coniferous woodland

     G3.1          [Abies] and [Picea] woodland

!    G3.15        Southern Apennine [Abies alba] forests

!    G3.16        Moesian [Abies alba] forests

!    G3.17        Balkano-Pontic [Abies] forests

!    G3.19        [Abies pinsapo] forests

!    G3.1B        Alpine and Carpathian subalpine [Picea] forests

!    G3.1C        Inner range montane [Picea] forests

!    G3.1D       Hercynian subalpine [Picea] forests

     G3.1E        Southern European [Picea abies] forests

!    G3.1E1      Southeastern Moesian [Picea abies] forests

!    G3.1E3      Montenegrine [Picea abies] forests

!    G3.1E4      Pelagonide [Picea abies] forests

!    G3.1E5      Balkan Range [Picea abies] forests

!    G3.1G       [Picea omorika] forests

!    G3.1H       [Picea orientalis] forests

     G3.2          Alpine [Larix] - [Pinus cembra] woodland

!    G3.21        Eastern Alpine siliceous [Larix] and [Pinus cembra] forests

!    G3.22        Eastern Alpine calcicolous [Larix] and [Pinus cembra] forests

!    G3.25        Carpathian [Larix] and [Pinus cembra] forests

!    G3.26        [Larix polonica] forests

     G3.3          [Pinus uncinata] woodland

!    G3.31        [Pinus uncinata] forests with [Rhododendron ferrugineum]

!    G3.32        Xerocline [Pinus uncinata] forests

     G3.4          [Pinus sylvestris] woodland south of the taiga

!    G3.41        Caledonian forest

     G3.42        Middle European [Pinus sylvestris] forests

     G3.423      Western Eurasian steppe pine forests

!    G3.4232    Sarmatic steppe [Pinus sylvestris] forests

!    G3.4233    Carpathian steppe [Pinus sylvestris] woods

!    G3.4234    Pannonic steppe [Pinus sylvestris] woods

     G3.44        Spring heath [Pinus sylvestris] forests

!    G3.442      Carpathian relict calcicolous [Pinus sylvestris] forests

!    G3.4C        Southeastern European [Pinus sylvestris] forests

!    G3.4E        Ponto-Caucasian [Pinus sylvestris] forests

     G3.5          [Pinus nigra] woodland

!    G3.51        Alpino-Apennine [Pinus nigra] forests

!    G3.52        Western Balkanic [Pinus nigra] forests

!    G3.53        [Pinus salzmannii] forests

!    G3.54        Corsican [Pinus laricio] forests

!    G3.55        Calabrian [Pinus laricio] forests

!    G3.56        [Pinus pallasiana] and [Pinus banatica] forests

!    G3.6          Subalpine mediterranean [Pinus] woodland

     G3.7          Lowland to montane mediterranean [Pinus] woodland (excluding [Pinus nigra])

     G3.71        Maritime [Pinus pinaster ssp. atlantica] forests

!    G3.711      Charente [Pinus pinaster ssp. atlantica] - [Quercus ilex] forests

!    G3.712      Aquitanian [Pinus pinaster ssp. atlantica] - [Quercus suber] forests

!    G3.714      Iberian [Pinus pinaster ssp. atlantica] forests

!    G3.72        [Pinus pinaster ssp. pinaster] ([Pinus mesogeensis]) forests

!    G3.73        [Pinus pinea] forests

     G3.74        [Pinus halepensis] forests

!    G3.741      Iberian [Pinus halepensis] forests

!    G3.742      Balearic [Pinus halepensis] forests

!    G3.743      Provenço-Ligurian [Pinus halepensis] forests

!    G3.744      Corsican [Pinus halepensis] woods

!    G3.745      Sardinian [Pinus halepensis] woods

!    G3.746      Sicilian [Pinus halepensis] woods

     G3.747      Italic [Pinus halepensis] forests

!    G3.7471    Gargano [Pinus halepensis] forests

!    G3.7472    Metapontine [Pinus halepensis] forests

!    G3.7473    Umbrian [Pinus halepensis] forests

!    G3.748      Hellenic [Pinus halepensis] forests

!    G3.749      Illyrian [Pinus halepensis] forests

!    G3.74A     East Mediterranean [Pinus halepensis] forests

!    G3.75        [Pinus brutia] forests

!    G3.8          Canary Island [Pinus canariensis] woodland

!    G3.9          Coniferous woodland dominated by [Cupressaceae] or [Taxaceae]

                       includes the following subtypes separately listed in or split unit from the 1998 version:

G3.9C     [Cedrus] woodland

!    G3.D         Boreal bog conifer woodland

!    G3.E          Nemoral bog conifer woodland

     H               Inland unvegetated or sparsely vegetated habitats

!    H1             Terrestrial underground caves, cave systems, passages and waterbodies

     H2             Screes

     H2.6          Calcareous and ultra-basic screes of warm exposures

     H2.61        Peri-Alpine thermophilous screes

!    H2.613      Paris Basin screes

     X               Habitat complexes

!    X01           Estuaries

!    X02           Saline coastal lagoons

!    X03           Brackish coastal lagoons

!    X04           Raised bog complexes

!    X18           Wooded steppe

!    X29           Salt lake islands

!    X35           New EUNIS complex ! "Inland Sand Dunes"